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| LUCANIDAE |
| AESALINAE |
| SYNDESINAE |
| DORCINAE |
| LUCANINAE |
|
|
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|
|
 |
| HYSOBORIDAE |
 |
| OCHODAEIDAE |
|
|
|
|
| CHIRONINAE |
| AEGIALIINAE |
| APHODIINAE |
| SCARABAEINAE |
| ORPHNINAE |
| MELOLONTHINAE |
| RUTELINAE |
| PACHYPODINAE |
| DYNASTINAE |
| CETONIINAE |
